Publications

Wang, B., Shi, G., Xu, C., Spicer, R.A., Perrichot, V., Schmidt, A.R., Feldberg, K., Heinrichs, J., Chny, C., Pang, H., Liu, X., Gao, T., Wang, Z., Slipinski, A., Solrzano-Kraemer, M.M., Heads, S.W., Thomas, M.J., Sadowski, E., Szwedo, J., Azar, D., Nel, A., Liu, Y., Chen, J., Zhang, Q., Zhang, Q., Luo, C., Yu, T., Zheng, D., Zhang, H., Engel, M.S. The mid-Miocene Zhangpu biota reveals an outstandingly rich rainforest biome in East Asia. Science Advances, 7(18) eabg0625, 30 April, 2021 DOI: 10.1126/sciadv.abg0625

Thomas, M.J., Skejo, J., & Heads, S.W. The last batrachideine of Europe: A new genus and species of pygmy grasshopper (Orthoptera: Tetrigidae) from Eocene Baltic amber. Zootaxa, 16 October, 2019.

Heads, S.W., Thomas, M.J. & Wang, Y. 2018. A new genus and species of Elcanidae (Insecta: Orthoptera) from Cretaceous Burmese amber. Zootaxa 4527: 575–580.

Makarkin, V.N., Wedmann, S. & Heads, S.W. 2018. A systematic reappraisal of Araripeneuridae (Neuroptera: Myrmeleontoidea), with description of new species from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 84: 600–621.

Heads, S.W., Miller, A.N. & Crane, J.L. 2017. On the name of the oldest fossil mushroom. Mycological Progress 16: 11–12.

Swanson, D.R., Taylor, S.J. & Heads, S.W. 2017. Nine true bugs (Heteroptera) newly-discovered in Illinois. Zootaxa 4269: 571–585.

Makarkin, V.N., Heads, S.W. & Wedmann, S. 2017. Taxonomic study of the Cretaceous lacewing family Babinskaiidae (Neuroptera: Myrmeleontoidea: Nymphidoidae), with description of new taxa. Cretaceous Research 78: 149–160.

Heads, S.W., Miller, A.N., Crane J.L., Thomas M.J., Ruffatto D.M., Methven A.S., et al. (2017) The oldest fossil mushroom. PLoS ONE 12(6): e0178327. doi: https://doi.org/10.1371/journal.pone.0178327

Heads, S., & Hollier, J. 2016. The type specimen and generic placement of Tridactylus galla Saussure, 1895 (Orthoptera: Caelifera: Tridactylidae). Zootaxa 4193(1): 195–196. doi: https://dx.doi.org/10.11646/zootaxa.4193.1.12

Jepson, J., & Heads, S. 2016. Fossil Megaloptera (Insecta: Neuropterida) from the Lower Cretaceous Crato Formation of Brazil. Zootaxa 4098(1): 134–144. doi: https://dx.doi.org/10.11646/zootaxa.4098.1.5

Barling, N., Martill, D.M., Heads, S.W., Gallien, F. 2015. High fidelity preservation of fossil insects from the Crato Formation (Lower Cretaceous) of Brazil. Cretaceous Research 52(B): 605-622. doi: https://doi.org/10.1016/j.cretres.2014.05.007

Heads, S.W., Thomas, M.J., Wang, Y. 2014. A remarkable new pygmy grasshopper (Orthoptera, Tetrigidae) in Miocene amber from the Dominican Republic. ZooKeys 429: 87–100. doi:  https://dx.doi.org/10.3897/zookeys.429.8020

Barling, N., Heads, S.W. & Martill, D.M. 2013. A new parasitoid wasp (Hymenoptera: Chalcidoidea) from the Lower Cretaceous Crato Formation of Brazil: the first Mesozoic Pteromalidae. Cretaceous Research 45: 258–264. doi: https://doi.org/10.1016/j.cretres.2013.05.001

Heads, S.W. & Wang, Y. 2013. First fossil record of Melanoplus differentialis (Orthoptera: Acrididae: Melanoplinae). Entomological News 123: 33–37. doi: https://doi.org/10.3157/021.123.0108

Jepson, J.E., Heads, S.W., Makarkin, V.N. & Ren, D. 2013. New fossil mantidflies (Insecta, Neuroptera, Mantispidae) from the Mesozoic of northeastern China. Palaeontology 56: 603–613. doi: https://doi.org/10.1111/pala.12005

Heads, S.W., Penney, D. & Green, D.I. 2012. A new species of the cricket genus Proanaxipha in Miocene amber from the Dominican Republic (Orthoptera, Gryllidae, Pentacentrinae). ZooKeys 229: 111–118.

Pérez-de la Fuente, R., Heads, S.W., Hinojosa-Díaz, I.A. & Engel, M.S. 2012. The first record of Protogryllinae from the Jurassic of India (Orthoptera: Grylloidea). Journal of the Kansas Entomological Society 85: 53–58. doi: https://doi.org/10.2317/JKES111103.1

Heads, S.W. & Leuzinger, L. 2011. On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera: Ensifera). ZooKeys 77: 17–30.

Heads, S.W. 2010. New Tridactyloidea in Miocene amber from the Dominican Republic (Orthoptera: Caelifera). Annales de la Société Entomologique de France 46: 204–210. doi: https://doi.org/10.1080/00379271.2010.10697659

Heads, S.W. 2010. The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae): 20 million years of troglobiomorphosis or exaptation in the dark? Zoological Journal of the Linnean Society 158: 56–65. doi: https://doi.org/10.1111/j.1096-3642.2009.00587.x

Heads, S.W. 2009. A new pygmy mole cricket in Cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia 26: 75–82. doi: https://doi.org/10.1016/j.cretres.2020.104371

Heads, S.W. 2009. New pygmy grasshoppers in Miocene amber from the Dominican Republic (Orthoptera: Tetrigidae). Denisia 26: 69–74.

Delclòs, X., Nel, A., Azar, D. Bechly, G., Dunlop, J.A., Engel, M.S. & Heads, S.W. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 247: 353–381.

Heads, S.W., Martill, D.M. & Loveridge, R.F. 2008. Palaeoentomological paradise: the Cretaceous Crato Formation of Brazil. Antenna 32: 91–98.

Heads, S.W. 2008. A new species of Yuripopovia (Coleorrhyncha: Progonocimicidae) from the Early Cretaceous of the Isle of Wight. British Journal of Entomology & Natural History 21: 247–254.

Heads, S.W. 2008. The first fossil Proscopiidae (Insecta, Orthoptera, Eumastacoidea) with comments on the historical biogeography and evolution of the family. Palaeontology 51: 499–507. doi: https://doi.org/10.1111/j.1475-4983.2008.00756.x

Martill, D.M., Bechly, G. & Heads, S.W. 2007. Appendix: species list for the Crato Formation, pp. 582–607 in: Martill, D.M., Bechly G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp.

Menon, F., Heads, S.W. & Szwedo, J. 2007. Cicadomorpha: cicadas and relatives, pp. 283–297 in: Martill, D.M., Bechly G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp.

Martins-Neto, R.G., Heads, S.W. & Bechly, G. 2007. Neuropterida: snakeflies, dobsonflies and lacewings, pp. 328–340 in: Martill, D.M., Bechly G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp.

Heads, S.W. & Martins-Neto, R.G. 2007. Orthopterida; grasshoppers, crickets, locusts and stick insects, pp. 265–283 in: Martill, D.M., Bechly G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp.

Heads, S.W. 2006 [for 2005]. A new caddisfly larval case (Insecta, Trichoptera) from the Lower Cretaceous Vectis Formation (Wealden Group) of the Isle of Wight, southern England. Proceedings of the Geologists’ Association 117: 307–310. doi: https://doi.org/10.1016/S0016-7878(06)80037-4

Menon, F., Heads, S.W. & Martill, D.M. 2005. New Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 26: 837–844. doi: https://doi.org/10.1016/j.cretres.2005.05.005

Menon, F. & Heads, S.W. 2005. New species of Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) 357: 1–11. (PDF)

Heads, S.W., Martill, D.M. & Loveridge, R.F. 2005. An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology 48: 1409–1417.  doi: 10.1111/j.1475-4983.2005.00514.x

Posters

Mierzejewski, C.M., Thomas, M.J., Heads, S.W. 2017. Exceptionally Preserved Orthoptera (Arthropoda: Insecta) from the Late Pennsylvanian Kinney Brick Quarry Lagerstatte of New Mexico. (PDF)

Nguy, W.N., Wittmer, J.M., Heads, S.W., Thomas, M.J. 2015. Rediscovery, Curation and Characterization of a Miocene rhinoceros (Perissodactlya: Rhinocerotidae ) hash from agate springs, Nebraska. 2015 SESE Research Review. (PDF)

Thomas, M.J., Nicholson, S.H., Best, M., Heads, S.W. 2014. The Milton Sanderson Dominican Amber Collection at the INHS. 2014 Prairie Lightning Symposium Poster Presentation. (PDF)